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The two stations share studio facilities at the Fox Television Center in West Los Angeles; KCOP's transmitter is located atop Mount Wilson. It was originally co-owned with local radio station KMTR (now KLAC).
Operating as an independent station early on, it began running some programming from the Du Mont Television Network in 1949 after KTLA (channel 5) disaffiliated from the network after a one-year tenure.
On February 22, 2006, News Corporation announced the launch of a new "sixth" network called My Network TV, which would have KCOP and the other Fox-owned UPN stations (plus an independent station in Dallas–Fort Worth) serve as the nuclei.
UPN continued to broadcast on stations across the country until September 15, 2006.
Rumors persisted that UPN would move to the higher-rated KCAL, reverting KCOP to independent station status.
During the late 1990s, the station began carrying a large amount of younger leaning talk shows (such as The Ricki Lake Show, The Jenny Jones Show, and The Montel Williams Show), reality series, some sitcoms during the evening hours, and syndicated cartoons (such as Double Dragon) in the morning well as the popular anime series Sailor Moon.